Faculty/Staff Directory

First Name

Last Name

Profile Photo


Assistant Professor

Office Address
3313 Kindschi Hall of Science

Office Phone

Areas of Expertise
Phycology, Barcoding, Phylogenetics, Taxonomy, Stream Ecology

Courses Taught at GVSU

BIO 121 General Biology II
BIO 413 Freshwater Algae
BIO 440 Limnology

Research Interests

Aquatic ecology
Biological indicators
Effects of UV radiation
Evolution of chloroplast and mitochondrial genomes
Algal biogeography
Diatom taxonomy and systematics
Using molecular tools to answer biological questions


Assistant Research Professor, Temple University, Biology, 2017-2018.
Post-doctoral Fellow, University of Colorado, Boulder, Museum of Natural History, 2012-2013.
Ph.D., University of New Brunswick, Fredericton, Biology, 2007-2012.
Diploma in University Teaching, University of New Brunswick, Fredericton, 2010.
M.S., Ohio University, Environmental and Plant Biology, 2001-2003.
B.S., Juniata College, Writing and the Environment, 1995-1999.

Select Publications

Pogoda CS, Keepers KG, Hamsher SE, Stepanek JG, Kane NC, Kociolek JP. 2018. Comparative analysis of the mitochondrial genomes of six newly sequenced diatoms reveals group II introns in the barcoding region of cox1. Mitochondrial DNA Part A, doi: 10.1080/24701394.2018.1450397

Hamsher SE, Sung CYT, Sanders RW. 2017. Effects of temperature and photorepair radiation on a marine ciliate exposed to UVB radiation. Journal of Eukaryotic Microbiology. 65: 458–467.

Kociolek JP, Hamsher SE, Kulikovskiy M, Bramburger AJ. 2017. Are there species flocks in freshwater diatoms? A review of past reports and a look to the future. Hydrobiologia. 792: 17–35.

Hamsher SE, Graeff CL, Stepanek JG, Kociolek JP. 2014. Frustular morphology and polyphyly in freshwater Denticula (Bacillariophyta) species, and the description of Tetralunata gen. nov. (Epithemiaceae, Rhopalodiales). Plant Ecology and Evolution. 147: 346–365.

Hamsher SE, Evans KM, Mann DG, Poulickova A, Saunders GW. 2011. Barcoding diatoms: exploring alternatives to COI-5P. Protist. 162: 405–422.

Hamsher SE, Verb RG, Vis ML. 2004. Analysis of acid mine drainage impacted streams using a periphyton index. Journal of Freshwater Ecology. 19: 313–324.


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